Vertebrate footprints from the Kem Kem beds (Morocco): A novel ichnological approach to faunal reconstruction
The first survey of vertebrate fossil tracks and trackways from the well-known Kem Kem beds of southern Morocco is here presented. The aims are to: (1) discuss the diversity of vertebrate trackmakers represented in two trace-yielding units at three localities, and (2) apply the novel data to ongoing...
Saved in:
| Published in | Palaeogeography, palaeoclimatology, palaeoecology Vol. 383-384; pp. 52 - 58 |
|---|---|
| Main Authors | , , , , , , |
| Format | Journal Article |
| Language | English |
| Published |
Elsevier B.V
01.08.2013
|
| Subjects | |
| Online Access | Get full text |
Cover
Loading…
| Abstract | The first survey of vertebrate fossil tracks and trackways from the well-known Kem Kem beds of southern Morocco is here presented. The aims are to: (1) discuss the diversity of vertebrate trackmakers represented in two trace-yielding units at three localities, and (2) apply the novel data to ongoing debates about vertebrate ecological associations within the Kem Kem. The majority of the tracks we recorded from the Kem Kem are tridactyl theropod footprints; other dinosaurian track records include those of possible ornithopod dinosaurs, extremely rare as body fossils within Kem Kem collections. Traces of swimming turtles are reported, alongside the first tracks of crocodyliforms and possible pterosaurs to be recorded from the Moroccan Cretaceous (both unsurprising given their abundance as body fossil records in this region). Differences between the collected ichnological sample and faunal reconstructions made on the basis of skeletal evidence are discussed; as trace fossils record the same environment in which the trackmaker lived (i.e. not subject to post-mortem transportation), data of this type arguably provide a more precise palaeoecological sample than the heavily re-worked and usually fragmentary body fossils from the Kem Kem.
•Report and analysis of several vertebrate traces in the Cretaceous “Kem Kem beds”•First occurrence of ornithopod, crocodyliform, and pterosaur tracks in the area•Use of trace assemblage for palaeofaunal reconstructions•Differences with the bone fauna as a key for environmental consideration |
|---|---|
| AbstractList | The first survey of vertebrate fossil tracks and trackways from the well-known Kem Kem beds of southern Morocco is here presented. The aims are to: (1) discuss the diversity of vertebrate trackmakers represented in two trace-yielding units at three localities, and (2) apply the novel data to ongoing debates about vertebrate ecological associations within the Kem Kem. The majority of the tracks we recorded from the Kem Kem are tridactyl theropod footprints; other dinosaurian track records include those of possible ornithopod dinosaurs, extremely rare as body fossils within Kem Kem collections. Traces of swimming turtles are reported, alongside the first tracks of crocodyliforms and possible pterosaurs to be recorded from the Moroccan Cretaceous (both unsurprising given their abundance as body fossil records in this region). Differences between the collected ichnological sample and faunal reconstructions made on the basis of skeletal evidence are discussed; as trace fossils record the same environment in which the trackmaker lived (i.e. not subject to post-mortem transportation), data of this type arguably provide a more precise palaeoecological sample than the heavily re-worked and usually fragmentary body fossils from the Kem Kem. The first survey of vertebrate fossil tracks and trackways from the well-known Kem Kem beds of southern Morocco is here presented. The aims are to: (1) discuss the diversity of vertebrate trackmakers represented in two trace-yielding units at three localities, and (2) apply the novel data to ongoing debates about vertebrate ecological associations within the Kem Kem. The majority of the tracks we recorded from the Kem Kem are tridactyl theropod footprints; other dinosaurian track records include those of possible ornithopod dinosaurs, extremely rare as body fossils within Kem Kem collections. Traces of swimming turtles are reported, alongside the first tracks of crocodyliforms and possible pterosaurs to be recorded from the Moroccan Cretaceous (both unsurprising given their abundance as body fossil records in this region). Differences between the collected ichnological sample and faunal reconstructions made on the basis of skeletal evidence are discussed; as trace fossils record the same environment in which the trackmaker lived (i.e. not subject to post-mortem transportation), data of this type arguably provide a more precise palaeoecological sample than the heavily re-worked and usually fragmentary body fossils from the Kem Kem. •Report and analysis of several vertebrate traces in the Cretaceous “Kem Kem beds”•First occurrence of ornithopod, crocodyliform, and pterosaur tracks in the area•Use of trace assemblage for palaeofaunal reconstructions•Differences with the bone fauna as a key for environmental consideration |
| Author | Dyke, Gareth J. Breda, Anna Belvedere, Matteo Jalil, Nour-Eddine Bourget, Hélène Gattolin, Giovanni Khaldoune, Fatima |
| Author_xml | – sequence: 1 givenname: Matteo surname: Belvedere fullname: Belvedere, Matteo email: matteo.belvedere@hotmail.com organization: Ocean and Earth Sciences, University of Southampton, European Way, Southampton SO14 3ZH, United Kingdom – sequence: 2 givenname: Nour-Eddine surname: Jalil fullname: Jalil, Nour-Eddine email: nourjalil@yahoo.fr, njalil@ucam.ac.ma organization: Department of Earth Sciences - Vertebrate Evolution and Palaeoenvironments, Université Cadi Ayyad, Faculty of Sciences Semlaia, Prince Moulay Abdellah Boulevard, P.O. Box 2390, Marrakech, Morocco – sequence: 3 givenname: Anna surname: Breda fullname: Breda, Anna email: anna.breda@unipd.it organization: Department of Geosciences, University of Padova, Via G. Gradenigo 6, 35131 Padova, Italy – sequence: 4 givenname: Giovanni surname: Gattolin fullname: Gattolin, Giovanni email: giovanni.gattolin@studenti.unipd.it organization: Department of Geosciences, University of Padova, Via G. Gradenigo 6, 35131 Padova, Italy – sequence: 5 givenname: Hélène surname: Bourget fullname: Bourget, Hélène email: helene@yahoo.fr organization: Museum of Natural History of Marrakesh, Allah El Fassi Boulevard, Marrakech, Morocco – sequence: 6 givenname: Fatima surname: Khaldoune fullname: Khaldoune, Fatima email: khaldoune_fa@yahoo.fr organization: Department of Earth Sciences - Vertebrate Evolution and Palaeoenvironments, Université Cadi Ayyad, Faculty of Sciences Semlaia, Prince Moulay Abdellah Boulevard, P.O. Box 2390, Marrakech, Morocco – sequence: 7 givenname: Gareth J. surname: Dyke fullname: Dyke, Gareth J. email: gareth.dyke@soton.ac.uk organization: Ocean and Earth Sciences, University of Southampton, European Way, Southampton SO14 3ZH, United Kingdom |
| BookMark | eNqFUU1v1DAQtVCR2C78Aw4-lkPC2E7ipAekqiofoogLIG6W44xZrxLPYnsr8e_Jspw40MNopJn3RvPeu2QXkSIy9lJALUB0r_f1wc4WqZYgVA1NDbJ7wjai17LqRPf9gm0AlKhA9PIZu8x5D7BClNyw5RumgmOyBbknKocUYsncJ1p42SH_iMufGnHK_OoTJXKOXl3zGx7pAWce3C7STD-CszO3h0Mi63a8EPf2GNdRQkcxl3R0JVB8zp56O2d88bdv2de3d19u31f3n999uL25r6wa2lJJ2w_YN-i11aharScFvuug1w14DyD81Avn_Ogb7TU0Qul27GGcWuXUulBbdnW-u_7z84i5mCVkh_NsI9IxG7nqV7LtBv0oVHRSD33bDO0KvT5DXaKcE3rjQrEnXSXZMBsB5hSH2ZtzHOYUh4HGnLzesuYf8mr1YtOvx2hvzjRc7XoImEx2AaPDKazWFjNR-P-B31bkqTw |
| CitedBy_id | crossref_primary_10_1016_j_palaeo_2019_109394 crossref_primary_10_1371_journal_pone_0115477 crossref_primary_10_1016_j_cretres_2015_02_002 crossref_primary_10_1016_j_jafrearsci_2017_06_013 crossref_primary_10_1016_j_palaeo_2024_112218 crossref_primary_10_1080_10420940_2017_1337010 crossref_primary_10_1016_j_palaeo_2024_112338 crossref_primary_10_1016_j_geobios_2019_12_002 crossref_primary_10_1016_j_jafrearsci_2019_103547 crossref_primary_10_1371_journal_pone_0180289 crossref_primary_10_1016_j_cretres_2017_12_011 crossref_primary_10_1080_02724634_2024_2311791 crossref_primary_10_1080_10420940_2016_1265524 crossref_primary_10_7717_peerj_1323 crossref_primary_10_1111_bij_12308 crossref_primary_10_1144_SP545_2023_141 crossref_primary_10_1016_j_jafrearsci_2024_105306 crossref_primary_10_1017_S0016756815000576 crossref_primary_10_7717_peerj_11544 crossref_primary_10_1093_zoolinnean_zlaa166 crossref_primary_10_1016_j_palaeo_2017_11_025 crossref_primary_10_1016_j_palaeo_2017_01_050 crossref_primary_10_1080_08912963_2023_2269173 crossref_primary_10_1002_ar_25024 crossref_primary_10_1371_journal_pone_0144695 crossref_primary_10_1016_j_palaeo_2018_02_028 crossref_primary_10_1007_s12542_025_00716_1 crossref_primary_10_1111_bij_12340 crossref_primary_10_1016_j_pgeola_2013_09_003 crossref_primary_10_1080_10420940_2023_2258261 crossref_primary_10_1016_j_jsames_2020_102936 crossref_primary_10_1111_jzo_12110 crossref_primary_10_1007_s41513_023_00212_y crossref_primary_10_3390_d14111007 crossref_primary_10_1080_10420940_2020_1763337 crossref_primary_10_1016_j_jafrearsci_2017_07_004 crossref_primary_10_1016_j_earscirev_2020_103321 crossref_primary_10_3989_egeol_41821_341 crossref_primary_10_1016_j_cretres_2022_105310 crossref_primary_10_1111_pala_12132 crossref_primary_10_1093_zoolinnean_zlae109 crossref_primary_10_7717_peerj_5925 crossref_primary_10_3897_zookeys_928_47517 crossref_primary_10_1080_10420940_2017_1386661 crossref_primary_10_1016_j_cretres_2022_105388 crossref_primary_10_1016_j_cretres_2024_105987 crossref_primary_10_1016_j_eve_2023_100006 crossref_primary_10_1007_s12542_022_00642_6 crossref_primary_10_1371_journal_pone_0090751 |
| Cites_doi | 10.1080/10420940601010943 10.1016/j.jafrearsci.2009.12.007 10.1126/science.272.5264.986 10.1016/j.cub.2010.10.001 10.22179/REVMACN.7.333 10.1006/cres.2001.0276 10.2475/ajs.293.A.167 10.4202/app.2011.0043 10.1371/journal.pone.0010875 10.3897/zookeys.28.325 10.1130/G30188A.1 10.1080/10420940109380188 10.1371/journal.pone.0030471 10.1080/10420940601006826 10.1016/j.palaeo.2013.04.011 |
| ContentType | Journal Article |
| Copyright | 2013 Elsevier B.V. |
| Copyright_xml | – notice: 2013 Elsevier B.V. |
| DBID | AAYXX CITATION 7SN 7ST C1K SOI 7S9 L.6 |
| DOI | 10.1016/j.palaeo.2013.04.026 |
| DatabaseName | CrossRef Ecology Abstracts Environment Abstracts Environmental Sciences and Pollution Management Environment Abstracts AGRICOLA AGRICOLA - Academic |
| DatabaseTitle | CrossRef Ecology Abstracts Environment Abstracts Environmental Sciences and Pollution Management AGRICOLA AGRICOLA - Academic |
| DatabaseTitleList | AGRICOLA Ecology Abstracts |
| DeliveryMethod | fulltext_linktorsrc |
| Discipline | Geology |
| EISSN | 1872-616X |
| EndPage | 58 |
| ExternalDocumentID | 10_1016_j_palaeo_2013_04_026 S0031018213002137 |
| GeographicLocations | Morocco |
| GeographicLocations_xml | – name: Morocco |
| GroupedDBID | --K --M -DZ -~X .~1 0R~ 123 1B1 1RT 1~. 1~5 29O 4.4 457 4G. 53G 5VS 7-5 71M 8P~ 9JM 9JN AABNK AACTN AAEDT AAEDW AAIAV AAIKJ AAKOC AALRI AAOAW AAQFI AAQXK AAXUO ABFNM ABFYP ABLJU ABLST ABMAC ABQEM ABQYD ABXDB ABYKQ ACDAQ ACGFS ACLVX ACRLP ACSBN ADBBV ADEZE ADKLL ADMUD AEBSH AEKER AENEX AFKWA AFMIJ AFTJW AFXIZ AGHFR AGUBO AGYEJ AHEUO AHHHB AIEXJ AIKHN AITUG AJBFU AJOXV AKIFW ALMA_UNASSIGNED_HOLDINGS AMFUW AMRAJ ASPBG ATOGT AVWKF AXJTR AZFZN BKOJK BLECG BLXMC CS3 DU5 EBS EFJIC EFLBG EJD EO8 EO9 EP2 EP3 F5P FDB FEDTE FGOYB FIRID FNPLU FYGXN G-2 G-Q GBLVA HMA HVGLF HZ~ IHE IMUCA J1W KCYFY KOM LY3 M41 MO0 MVM N9A O-L O9- OAUVE OZT P-8 P-9 P2P PC. Q38 R2- RIG ROL RPZ SCC SDF SDG SDP SEP SES SEW SPC SPCBC SSE SSJ SSZ T5K TN5 WUQ XJT XOL XPP ZCG ZMT ~02 ~G- AAHBH AATTM AAXKI AAYWO AAYXX ABWVN ACRPL ACVFH ADCNI ADNMO ADXHL AEIPS AEUPX AFJKZ AFPUW AGCQF AGQPQ AGRNS AIGII AIIUN AKBMS AKRWK AKYEP ANKPU APXCP BNPGV CITATION SSH 7SN 7ST C1K SOI 7S9 EFKBS L.6 |
| ID | FETCH-LOGICAL-a395t-2a89e84ef7a7e3577d30f6608740ff001fd81ccfbf47f7041375b80bd53c31cc3 |
| IEDL.DBID | .~1 |
| ISSN | 0031-0182 |
| IngestDate | Tue Aug 05 09:50:01 EDT 2025 Thu Jul 10 18:30:55 EDT 2025 Tue Jul 01 03:15:58 EDT 2025 Thu Apr 24 22:56:44 EDT 2025 Fri Feb 23 02:32:53 EST 2024 |
| IsPeerReviewed | true |
| IsScholarly | true |
| Keywords | Vertebrate ichnology Palaeoecology Kem Kem beds Morocco Cretaceous |
| Language | English |
| LinkModel | DirectLink |
| MergedId | FETCHMERGED-LOGICAL-a395t-2a89e84ef7a7e3577d30f6608740ff001fd81ccfbf47f7041375b80bd53c31cc3 |
| Notes | ObjectType-Article-1 SourceType-Scholarly Journals-1 ObjectType-Feature-2 content type line 23 |
| PQID | 1627985495 |
| PQPubID | 23462 |
| PageCount | 7 |
| ParticipantIDs | proquest_miscellaneous_2000325697 proquest_miscellaneous_1627985495 crossref_citationtrail_10_1016_j_palaeo_2013_04_026 crossref_primary_10_1016_j_palaeo_2013_04_026 elsevier_sciencedirect_doi_10_1016_j_palaeo_2013_04_026 |
| ProviderPackageCode | CITATION AAYXX |
| PublicationCentury | 2000 |
| PublicationDate | August 2013 2013-08-00 20130801 |
| PublicationDateYYYYMMDD | 2013-08-01 |
| PublicationDate_xml | – month: 08 year: 2013 text: August 2013 |
| PublicationDecade | 2010 |
| PublicationTitle | Palaeogeography, palaeoclimatology, palaeoecology |
| PublicationYear | 2013 |
| Publisher | Elsevier B.V |
| Publisher_xml | – name: Elsevier B.V |
| References | Depéret, Savornin (bb0045) 1925; 181 Avanzini, García-Ramos, Lires, Menegon, Piñuela, Fernández (bb0165) 2005; 50 Lockley, Harris, Mitchell (bb9005) 2008; 28 Amiot, Buffetaut, Tong, Boudad, Kabiri (bb0005) 2004; 9 Dyke (bb0050) 2010; 20 Marty (bb0110) 2008; 21 Calvo, Lockley (bb9010) 2001; 22 Lockley, Meyer, Santos (bb0100) 2000; 15 Avanzini, García-Ramos, Lires, Piñuela, Lockley (bb0015) 2007; 14 McGowan, Dyke (bb0120) 2009; 37 Lavocat (bb0090) 1954; 1 Marty, Meyer, Paratte, Lovis, Cattin (bb0115) 2012 Sereno, Larsson (bb0140) 2009; 28 Hunt, Lucas (bb0070) 2007; 14 Läng, Boudad, Maio, Samankassou, Tabouelle, Haiyan, Cavin (bb0085) July 2013; 381–382 Lockley, Wright (bb0105) 2001 Leonardi (bb0095) 1987 Thulborn (bb0155) 2001; 8 Jalil, Belvedere, Breda, Gattolin, Dyke (bb0175) 2011; 3 Cau, Dalla Vecchia, Fabbri (bb0030) 2012; 57 Cavin, Tong, Boudad, Meister, Piuz, Tabouelle, Aarab, Amiot, Buffetaut, Dyke (bb0170) 2010; 57 Farlow, Chapman (bb0060) 1997 Holliday, Gardner (bb0065) 2012; 7 Weishampel, Dodson, Osmólska (bb0160) 2004 Russell (bb0130) 1996; 18 Sereno, Dutheil, Iarochene, Larsson, Lyon, Magwene, Sidor, Varricchio, Wilson (bb0135) 1996; 272 Belvedere, Mietto, Ishigaki (bb0020) 2010; 54 Farlow (bb0055) 1993; 293A Novas, Dalla Vecchia, Pais (bb0125) 2005; 7 Rodrigues, Kellner, Mader, Russell (bb9000) 2011; 117 D'Orazi Porchetti, Nicosia, Biava, Maganuco (bb0040) 2011; 177 Thulborn (bb0150) 1990 Ibrahim, Unwin, Martill, Baidder, Zouhri (bb0075) 2010; 5 Cau (10.1016/j.palaeo.2013.04.026_bb0030) 2012; 57 D'Orazi Porchetti (10.1016/j.palaeo.2013.04.026_bb0040) 2011; 177 Leonardi (10.1016/j.palaeo.2013.04.026_bb0095) 1987 Marty (10.1016/j.palaeo.2013.04.026_bb0115) 2012 Holliday (10.1016/j.palaeo.2013.04.026_bb0065) 2012; 7 Sereno (10.1016/j.palaeo.2013.04.026_bb0140) 2009; 28 Belvedere (10.1016/j.palaeo.2013.04.026_bb0020) 2010; 54 Amiot (10.1016/j.palaeo.2013.04.026_bb0005) 2004; 9 Avanzini (10.1016/j.palaeo.2013.04.026_bb0165) 2005; 50 Lockley (10.1016/j.palaeo.2013.04.026_bb0105) 2001 Ibrahim (10.1016/j.palaeo.2013.04.026_bb0075) 2010; 5 Thulborn (10.1016/j.palaeo.2013.04.026_bb0150) 1990 Lavocat (10.1016/j.palaeo.2013.04.026_bb0090) 1954; 1 Russell (10.1016/j.palaeo.2013.04.026_bb0130) 1996; 18 Avanzini (10.1016/j.palaeo.2013.04.026_bb0015) 2007; 14 Lockley (10.1016/j.palaeo.2013.04.026_bb9005) 2008; 28 Marty (10.1016/j.palaeo.2013.04.026_bb0110) 2008; 21 Novas (10.1016/j.palaeo.2013.04.026_bb0125) 2005; 7 Calvo (10.1016/j.palaeo.2013.04.026_bb9010) 2001; 22 Lockley (10.1016/j.palaeo.2013.04.026_bb0100) 2000; 15 Farlow (10.1016/j.palaeo.2013.04.026_bb0060) 1997 Hunt (10.1016/j.palaeo.2013.04.026_bb0070) 2007; 14 Sereno (10.1016/j.palaeo.2013.04.026_bb0135) 1996; 272 Rodrigues (10.1016/j.palaeo.2013.04.026_bb9000) 2011; 117 Läng (10.1016/j.palaeo.2013.04.026_bb0085) 2013; 381–382 Cavin (10.1016/j.palaeo.2013.04.026_bb0170) 2010; 57 McGowan (10.1016/j.palaeo.2013.04.026_bb0120) 2009; 37 Thulborn (10.1016/j.palaeo.2013.04.026_bb0155) 2001; 8 Farlow (10.1016/j.palaeo.2013.04.026_bb0055) 1993; 293A Weishampel (10.1016/j.palaeo.2013.04.026_bb0160) 2004 Depéret (10.1016/j.palaeo.2013.04.026_bb0045) 1925; 181 Jalil (10.1016/j.palaeo.2013.04.026_bb0175) 2011; 3 Dyke (10.1016/j.palaeo.2013.04.026_bb0050) 2010; 20 |
| References_xml | – volume: 293A start-page: 167 year: 1993 end-page: 199 ident: bb0055 article-title: On the rareness of big, fierce animals: speculations about the body sizes, population densities, and geographic ranges of predatory mammals and large carnivorous dinosaurs publication-title: American Journal of Science – volume: 21 start-page: 1 year: 2008 end-page: 278 ident: bb0110 article-title: Sedimentology, taphonomy, and ichnology of Late Jurassic dinosaur tracks from the Jura carbonate platform (Chevenez—CombeRonde tracksite, NW Switzerland): insights into the tidal-flat palaeoenvironment and dinosaur diversity, locomotion, and palaeoecology publication-title: GeoFocus – volume: 3 start-page: 167 year: 2011 ident: bb0175 article-title: A varied ichnocoenosis in the Cretaceous Kem Kem beds of Morocco. 71st meeting of the Society of Vertebrate Paleontology, Abstract Book, Las Vegas, Nevada, USA, 2–5.11.2011 publication-title: Journal of Vertebrate Paleontology – volume: 117 start-page: 149 year: 2011 end-page: 160 ident: bb9000 article-title: New pterosaur specimens from the Kem Kem Beds (Upper Cretaceous, Cenomanian) of Morocco: Rivista Italiana di Paleontologia e Stratigrafia – volume: 9 start-page: 143 year: 2004 end-page: 149 ident: bb0005 article-title: Isolated theropod teeth from the Cenomanian of Morocco and their palaebiogeographical significance publication-title: Revue de Paléobiologie – volume: 22 start-page: 585 year: 2001 end-page: 590 ident: bb9010 article-title: The first pterosaur tracks from Gondwana publication-title: Cretaceous Research – volume: 57 start-page: 391 year: 2010 end-page: 412 ident: bb0170 article-title: Vertebrate assemblages from the early Late Cretaceous of southeastern Morocco: an overview publication-title: Journal of African Earth Sciences – volume: 5 start-page: e10875 year: 2010 ident: bb0075 article-title: A New Pterosaur (Pterodactyloidea: Azhdarchidae) from the Upper Cretaceous of Morocco publication-title: PLoS One – volume: 7 start-page: e30471 year: 2012 ident: bb0065 article-title: A New Eusuchian crocodyliform with novel cranial integument and its significance for the origin and evolution of Crocodylia publication-title: PLoS One – volume: 20 start-page: R983 year: 2010 end-page: R985 ident: bb0050 article-title: Palaeoecology: different dinosaur ecologies in deep time? publication-title: Current Biology – volume: 7 start-page: 167 year: 2005 end-page: 175 ident: bb0125 article-title: Theropod pedal unguals from the Late Cretaceous (Cenomanian) of Morocco, Africa publication-title: Revista del Museo Argentino de Ciencias Naturales, Nueva Serie – volume: 8 start-page: 207 year: 2001 end-page: 222 ident: bb0155 article-title: History and nomenclature of the theropod dinosaur tracks publication-title: Ichnos – year: 2004 ident: bb0160 article-title: The Dinosauria – volume: 181 start-page: 1108 year: 1925 end-page: 1111 ident: bb0045 article-title: Sur la découverte d'une faune de Vertébrés albiens à Timimoun (Sahara occidental) publication-title: Comptes Rendus des Hebdomadaires des Seances de l'Academie des Sciences à Paris – volume: 57 start-page: 661 year: 2012 end-page: 665 ident: bb0030 article-title: Evidence of a new carcharodontosaurid from the Upper Cretaceous of Morocco publication-title: Acta Palaeontologica Polonica – volume: 177 start-page: 463 year: 2011 end-page: 472 ident: bb0040 article-title: New Abelisaurid material from the Upper Cretaceous (Cenomanian) of Morocco publication-title: Rivista Italiana di Paleontologia e Stratigrafia – volume: 18 start-page: 349 year: 1996 end-page: 402 ident: bb0130 article-title: Isolated dinosaur bones from the Middle Cretaceous of the Tafilalt, Morocco publication-title: Bulletin du Muséum National d'Histoire Naturelle – volume: 37 start-page: 843 year: 2009 end-page: 846 ident: bb0120 article-title: A surfeit of theropods in the Moroccan Late Cretaceous? Comparing diversity estimates from field data and fossil shops publication-title: Geology – volume: 28 start-page: 1 year: 2009 end-page: 143 ident: bb0140 article-title: Cretaceous crocodyliforms from the Sahara publication-title: ZooKeys – volume: 54 start-page: 367 year: 2010 end-page: 380 ident: bb0020 article-title: A Late Jurassic diverse ichnocoenosis from the siliciclastic Iouaridene Formation (Central High Atlas, Morocco) publication-title: Geological Quarterly – volume: 381–382 start-page: 26 year: July 2013 end-page: 32 ident: bb0085 article-title: Unbalanced food web in a Late Cretaceous dinosaur assemblage publication-title: Palaeogeography, Palaeoclimatology, Palaeoecology – volume: 14 start-page: 143 year: 2007 end-page: 153 ident: bb0015 article-title: Crocodylomorph tracks from the Late Jurassic of Asturias (Spain) publication-title: Ichnos – volume: 50 start-page: 743 year: 2005 end-page: 755 ident: bb0165 article-title: Turtle tracks from the Late Jurassic of Asturias, Spain publication-title: Acta Palaeontologica Polonica – start-page: 65 year: 2012 end-page: 68 ident: bb0115 article-title: Spatial relationships in dinosaur ichnoassemblages and its influence on paleobiological interpretations—a case study from the Late Jurassic of NW Switzerland publication-title: Qijiang International Dinosaur Track Symposium, Chongqing Municipality, China, 28–20.11.2012 – year: 1987 ident: bb0095 article-title: Glossary and Manual of Tetrapod Footprint Palaeoichnology: Brasil – volume: 1 start-page: 65 year: 1954 end-page: 98 ident: bb0090 article-title: Sur les dinosauriens du Continental Intercalaire des Kem-Kem de la Daoura publication-title: Comptes Rendus 19th International Geological Congress 1952 – volume: 272 start-page: 986 year: 1996 end-page: 991 ident: bb0135 article-title: Predatory dinosaurs from the Sahara and Late Cretaceous faunal differentiation publication-title: Science – volume: 14 start-page: 59 year: 2007 end-page: 68 ident: bb0070 article-title: Tetrapod ichnofacies: a new paradigm publication-title: Ichnos – start-page: 519 year: 1997 end-page: 553 ident: bb0060 article-title: The scientific study of dinosaur footprints publication-title: The Complete Dinosaur – volume: 15 start-page: 313 year: 2000 end-page: 337 ident: bb0100 article-title: and the problematic concept of publication-title: Gaia: Revista de Geocienciasn – start-page: 428 year: 2001 end-page: 442 ident: bb0105 article-title: Trackways of Large Quadrupedal Ornithopods from the Cretaceous: a review publication-title: Mesozoic Vertebrate Life – volume: 28 start-page: 187 year: 2008 end-page: 198 ident: bb9005 article-title: A global overview of pterosaur ichnology: tracksite distribution in space and time publication-title: Flugsaurier: Pterosaur Papers in Honour of Peter Wellnhofer – year: 1990 ident: bb0150 article-title: Dinosaur Tracks – volume: 14 start-page: 143 issue: 1–2 year: 2007 ident: 10.1016/j.palaeo.2013.04.026_bb0015 article-title: Crocodylomorph tracks from the Late Jurassic of Asturias (Spain) publication-title: Ichnos doi: 10.1080/10420940601010943 – start-page: 428 year: 2001 ident: 10.1016/j.palaeo.2013.04.026_bb0105 article-title: Trackways of Large Quadrupedal Ornithopods from the Cretaceous: a review – year: 1990 ident: 10.1016/j.palaeo.2013.04.026_bb0150 – year: 2004 ident: 10.1016/j.palaeo.2013.04.026_bb0160 – volume: 57 start-page: 391 issue: 5 year: 2010 ident: 10.1016/j.palaeo.2013.04.026_bb0170 article-title: Vertebrate assemblages from the early Late Cretaceous of southeastern Morocco: an overview publication-title: Journal of African Earth Sciences doi: 10.1016/j.jafrearsci.2009.12.007 – volume: 15 start-page: 313 year: 2000 ident: 10.1016/j.palaeo.2013.04.026_bb0100 article-title: Megalosauripus and the problematic concept of Megalosaur footprints publication-title: Gaia: Revista de Geocienciasn – volume: 272 start-page: 986 issue: 5264 year: 1996 ident: 10.1016/j.palaeo.2013.04.026_bb0135 article-title: Predatory dinosaurs from the Sahara and Late Cretaceous faunal differentiation publication-title: Science doi: 10.1126/science.272.5264.986 – volume: 20 start-page: R983 issue: 22 year: 2010 ident: 10.1016/j.palaeo.2013.04.026_bb0050 article-title: Palaeoecology: different dinosaur ecologies in deep time? publication-title: Current Biology doi: 10.1016/j.cub.2010.10.001 – volume: 50 start-page: 743 issue: 4 year: 2005 ident: 10.1016/j.palaeo.2013.04.026_bb0165 article-title: Turtle tracks from the Late Jurassic of Asturias, Spain publication-title: Acta Palaeontologica Polonica – volume: 7 start-page: 167 issue: 2 year: 2005 ident: 10.1016/j.palaeo.2013.04.026_bb0125 article-title: Theropod pedal unguals from the Late Cretaceous (Cenomanian) of Morocco, Africa publication-title: Revista del Museo Argentino de Ciencias Naturales, Nueva Serie doi: 10.22179/REVMACN.7.333 – volume: 22 start-page: 585 year: 2001 ident: 10.1016/j.palaeo.2013.04.026_bb9010 article-title: The first pterosaur tracks from Gondwana publication-title: Cretaceous Research doi: 10.1006/cres.2001.0276 – volume: 293A start-page: 167 year: 1993 ident: 10.1016/j.palaeo.2013.04.026_bb0055 article-title: On the rareness of big, fierce animals: speculations about the body sizes, population densities, and geographic ranges of predatory mammals and large carnivorous dinosaurs publication-title: American Journal of Science doi: 10.2475/ajs.293.A.167 – volume: 57 start-page: 661 issue: 3 year: 2012 ident: 10.1016/j.palaeo.2013.04.026_bb0030 article-title: Evidence of a new carcharodontosaurid from the Upper Cretaceous of Morocco publication-title: Acta Palaeontologica Polonica doi: 10.4202/app.2011.0043 – volume: 5 start-page: e10875 issue: 5 year: 2010 ident: 10.1016/j.palaeo.2013.04.026_bb0075 article-title: A New Pterosaur (Pterodactyloidea: Azhdarchidae) from the Upper Cretaceous of Morocco publication-title: PLoS One doi: 10.1371/journal.pone.0010875 – volume: 28 start-page: 1 year: 2009 ident: 10.1016/j.palaeo.2013.04.026_bb0140 article-title: Cretaceous crocodyliforms from the Sahara publication-title: ZooKeys doi: 10.3897/zookeys.28.325 – volume: 177 start-page: 463 issue: 3 year: 2011 ident: 10.1016/j.palaeo.2013.04.026_bb0040 article-title: New Abelisaurid material from the Upper Cretaceous (Cenomanian) of Morocco publication-title: Rivista Italiana di Paleontologia e Stratigrafia – volume: 37 start-page: 843 issue: 9 year: 2009 ident: 10.1016/j.palaeo.2013.04.026_bb0120 article-title: A surfeit of theropods in the Moroccan Late Cretaceous? Comparing diversity estimates from field data and fossil shops publication-title: Geology doi: 10.1130/G30188A.1 – volume: 8 start-page: 207 issue: 3-4 year: 2001 ident: 10.1016/j.palaeo.2013.04.026_bb0155 article-title: History and nomenclature of the theropod dinosaur tracks Bueckeburgichnus and Megalosauripus publication-title: Ichnos doi: 10.1080/10420940109380188 – volume: 7 start-page: e30471 issue: 1 year: 2012 ident: 10.1016/j.palaeo.2013.04.026_bb0065 article-title: A New Eusuchian crocodyliform with novel cranial integument and its significance for the origin and evolution of Crocodylia publication-title: PLoS One doi: 10.1371/journal.pone.0030471 – start-page: 519 year: 1997 ident: 10.1016/j.palaeo.2013.04.026_bb0060 article-title: The scientific study of dinosaur footprints – volume: 18 start-page: 349 issue: série 4 year: 1996 ident: 10.1016/j.palaeo.2013.04.026_bb0130 article-title: Isolated dinosaur bones from the Middle Cretaceous of the Tafilalt, Morocco publication-title: Bulletin du Muséum National d'Histoire Naturelle – volume: 14 start-page: 59 issue: 1–2 year: 2007 ident: 10.1016/j.palaeo.2013.04.026_bb0070 article-title: Tetrapod ichnofacies: a new paradigm publication-title: Ichnos doi: 10.1080/10420940601006826 – volume: 21 start-page: 1 year: 2008 ident: 10.1016/j.palaeo.2013.04.026_bb0110 article-title: Sedimentology, taphonomy, and ichnology of Late Jurassic dinosaur tracks from the Jura carbonate platform (Chevenez—CombeRonde tracksite, NW Switzerland): insights into the tidal-flat palaeoenvironment and dinosaur diversity, locomotion, and palaeoecology – start-page: 65 year: 2012 ident: 10.1016/j.palaeo.2013.04.026_bb0115 article-title: Spatial relationships in dinosaur ichnoassemblages and its influence on paleobiological interpretations—a case study from the Late Jurassic of NW Switzerland – year: 1987 ident: 10.1016/j.palaeo.2013.04.026_bb0095 – volume: 9 start-page: 143 year: 2004 ident: 10.1016/j.palaeo.2013.04.026_bb0005 article-title: Isolated theropod teeth from the Cenomanian of Morocco and their palaebiogeographical significance publication-title: Revue de Paléobiologie – volume: 181 start-page: 1108 year: 1925 ident: 10.1016/j.palaeo.2013.04.026_bb0045 article-title: Sur la découverte d'une faune de Vertébrés albiens à Timimoun (Sahara occidental) publication-title: Comptes Rendus des Hebdomadaires des Seances de l'Academie des Sciences à Paris – volume: 3 start-page: 167 year: 2011 ident: 10.1016/j.palaeo.2013.04.026_bb0175 article-title: A varied ichnocoenosis in the Cretaceous Kem Kem beds of Morocco. 71st meeting of the Society of Vertebrate Paleontology, Abstract Book, Las Vegas, Nevada, USA, 2–5.11.2011 publication-title: Journal of Vertebrate Paleontology – volume: 381–382 start-page: 26 year: 2013 ident: 10.1016/j.palaeo.2013.04.026_bb0085 article-title: Unbalanced food web in a Late Cretaceous dinosaur assemblage publication-title: Palaeogeography, Palaeoclimatology, Palaeoecology doi: 10.1016/j.palaeo.2013.04.011 – volume: 54 start-page: 367 issue: 3 year: 2010 ident: 10.1016/j.palaeo.2013.04.026_bb0020 article-title: A Late Jurassic diverse ichnocoenosis from the siliciclastic Iouaridene Formation (Central High Atlas, Morocco) publication-title: Geological Quarterly – volume: 1 start-page: 65 year: 1954 ident: 10.1016/j.palaeo.2013.04.026_bb0090 article-title: Sur les dinosauriens du Continental Intercalaire des Kem-Kem de la Daoura – volume: 28 start-page: 187 year: 2008 ident: 10.1016/j.palaeo.2013.04.026_bb9005 article-title: A global overview of pterosaur ichnology: tracksite distribution in space and time – volume: 117 start-page: 149 year: 2011 ident: 10.1016/j.palaeo.2013.04.026_bb9000 |
| SSID | ssj0002632 |
| Score | 2.3092742 |
| Snippet | The first survey of vertebrate fossil tracks and trackways from the well-known Kem Kem beds of southern Morocco is here presented. The aims are to: (1) discuss... |
| SourceID | proquest crossref elsevier |
| SourceType | Aggregation Database Enrichment Source Index Database Publisher |
| StartPage | 52 |
| SubjectTerms | Cretaceous dinosaurs fauna fossils Kem Kem beds Morocco Palaeoecology paleoecology surveys swimming turtles Vertebrate ichnology |
| Title | Vertebrate footprints from the Kem Kem beds (Morocco): A novel ichnological approach to faunal reconstruction |
| URI | https://dx.doi.org/10.1016/j.palaeo.2013.04.026 https://www.proquest.com/docview/1627985495 https://www.proquest.com/docview/2000325697 |
| Volume | 383-384 |
| hasFullText | 1 |
| inHoldings | 1 |
| isFullTextHit | |
| isPrint | |
| link | http://utb.summon.serialssolutions.com/2.0.0/link/0/eLvHCXMwnV1LS8QwEA6LIngRn_hcInjQQ7XbNE3qbRF1VfSk4i0kbQIru-3iVsGLv92ZtFUURfDQHtqEhpl0HpmZbwjZY0kepdplAchfdFB6UZByngfO8cQYmdjcwzVd3ySDu_jygT90yElbC4NplY3sr2W6l9bNk6OGmkeT4RBrfBnCTUUYkIEbVpTHscBdfvj2meaBeOQ1NCO6zTJqy-d8jtdEj7QvAewxD3iKEAs_q6dvgtprn7NFstCYjbRfr2yJdGyxTObOfVve1xUyvrdPFYaAK0tdWVZ4WldNKdaOULDw6JUd-8vYfEr3r0vQWll5cEz7tChf7IgOm_N15BhtYcZpVVKnn_G73m3-gJpdJXdnp7cng6BppBBolvIqiLRMrYytE1pYxoUABrgkCbEdn3OgqFwue1nmjIuFEyHoNcGNDE3OWcbgBVsjM0VZ2HVCwyzWRhhtOdghnFkptWRGxhqmcvDGNghr6aeyBmUcm12MVJtO9qhqqiukugpjBVTfIMHHrEmNsvHHeNGyRn3ZLQoUwR8zd1tOKviRMDqiC1s-T1UviUQqwV3mv4_BuiYGRmIqNv-9gi0yH_mWGphEuE1mgHd2BwybynT9zu2S2f7F1eDmHawc-KM |
| linkProvider | Elsevier |
| linkToHtml | http://utb.summon.serialssolutions.com/2.0.0/link/0/eLvHCXMwnV3LS_QwEA8-EL2ITz7fETzoodptmib1JqKuj_Wk4i0kbQL7sbaLWwUv_u3OpK2iKIKH9tAkNMwk80hmfkPIDkvyKNUuC0D-ooPSiYKU8zxwjifGyMTmHq6pd510b-OLe34_Ro7bXBgMq2xkfy3TvbRuvhw01DwY9vuY48sQbirCCxl4iXEyGcP2xTIG-68fcR4ISF5jM6LfLKM2f84HeQ31QPscwA7ziKeIsfC9fvoiqb36OZ0js43dSI_qqc2TMVsskKkzX5f3ZZE83NnHCu-AK0tdWVZ4XFeNKCaPUDDx6KV98I-x-Yju9kpQW1m5d0iPaFE-2wHtNwfsyDLa4ozTqqROP-F_vd_8jjW7RG5PT26Ou0FTSSHQLOVVEGmZWhlbJ7SwjAsBHHBJEmI9PudAU7lcdrLMGRcLJ0JQbIIbGZqcs4xBA1smE0VZ2H-EhlmsjTDacjBEOLNSasmMjDUM5eCOrRDW0k9lDcw4VrsYqDae7L-qqa6Q6iqMFVB9hQTvo4Y1zMYv_UXLGvVpuSjQBL-M3G45qWAn4fWILmz5NFKdJBKpBH-Z_9wHE5sYWImpWP3zDLbIdPemd6Wuzq8v18hM5OtrYEThOpkAPtoNsHIqs-lX8RtUrPox |
| openUrl | ctx_ver=Z39.88-2004&ctx_enc=info%3Aofi%2Fenc%3AUTF-8&rfr_id=info%3Asid%2Fsummon.serialssolutions.com&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.atitle=Vertebrate+footprints+from+the+Kem+Kem+beds+%28Morocco%29%3A+A+novel+ichnological+approach+to+faunal+reconstruction&rft.jtitle=Palaeogeography%2C+palaeoclimatology%2C+palaeoecology&rft.au=Belvedere%2C+Matteo&rft.au=Jalil%2C+Nour-Eddine&rft.au=Breda%2C+Anna&rft.au=Gattolin%2C+Giovanni&rft.date=2013-08-01&rft.issn=0031-0182&rft.volume=383&rft.spage=52&rft.epage=58&rft_id=info:doi/10.1016%2Fj.palaeo.2013.04.026&rft.externalDBID=NO_FULL_TEXT |
| thumbnail_l | http://covers-cdn.summon.serialssolutions.com/index.aspx?isbn=/lc.gif&issn=0031-0182&client=summon |
| thumbnail_m | http://covers-cdn.summon.serialssolutions.com/index.aspx?isbn=/mc.gif&issn=0031-0182&client=summon |
| thumbnail_s | http://covers-cdn.summon.serialssolutions.com/index.aspx?isbn=/sc.gif&issn=0031-0182&client=summon |